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"5. Conclusion
We have found, in both human and non-human animals, that darker pigmentation is associated with higher levels of aggression and sexuality (and in humans with lower IQ). Lighter pigmentation is associated with the slow reproductive strategy (K) including lower birth rates, less infant mortality, less violent crime, less HIV/AIDS, plus higher IQ, higher income, and greater longevity. The correlations between human pigmentation, aggression, and sexuality (and IQ), is further supported by the anthropological and sociological research on ‘‘pigmentocracies’’ (Lynn & Vanhanen, 2006). A pigmentocracy is a society in which status hierarchies are based largely on skin color, with lighter skin denoting higher status and darker skin lower status. Although these are typically explained by the legacy of slavery and imperialism, and although cultural and environmental factors undoubtedly play a substantial role (Rushton & Jensen, 2005), we have focused on genetic pleiotropy to explain the much less known relationship between skin
color and behavior.

Life history theory (LHT) may explain why darker individuals are more aggressive and sexually active and why these traits covary with longevity, birth rate, infant mortality, speed of maturation, and many other characteristics (Templer, 2008; Templer & Rushton, 2011). The melanocortin system is a physiological coordinator of pigmentation and life history traits. Skin color provides an important marker placing hormonal mediators such as testosterone in broader perspective. We recognize that this paper provides only a first approximation to what may become a workable explanation of melanin and its correlates. There are complex issues that need to be resolved. For example, skin coloration in humans is labile, showing much
unexplained variation such as the African Khoi-san in the Kalahari Desert who display light pigmentation. Nonetheless, we believe our data reviews point in a direction that could greatly advance knowledge. Although the traits do not always go in lockstep, our data and analyses raise new research directions that should be
seriously explored."

It's always interesting to see this stuff being explored, especially the more recent papers. Thanks for posting this.

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